Fred Stormshak
Distinguished Professor Emeritus
OSU Campus
Department of Animal and Rangeland Sciences
216 Withycombe Hall
Corvallis, OR 97331
United States


(1) Mechanisms of action of ovarian steroid hormones in regulating uterine gene expression as related to luteal function and embryonic mortality on the cow, ewe, and gilt. (2) Neuroendocrine basis for sexual behavior in sheep.


Ph.D. Endocrinology, University of Wisconsin, 1965

M.S. Washington State University, 1960

B.S. Washington State University, 1959


Roselli, C. and F. Stormshak. 2010. The ovine sexually dimorphic nucleus, aromatase, and sexual partner preference in sheep. J. Steroid Biochem. Mol. Biol. 118:252-256.

Roselli, C.E., C.T. Estill, H.L. Stadelman and F. Stormshak. 2009. The volume of the ovine sexually dimorphic nucleus of the preoptic area is independent of adult testosterone concentrations.  Brain Res. 1249:113-117.

Roselli, C., and F. Stormshak.  2009.  Prenatal programming of sexual partner preference:  the ram model.  J. Neuroendocrinol. 21:359-364.

Roselli, C., and F. Stormshak.  2009.  The neurobiology of sexual partner preferences in rams.  Horm. Behav. 55:611-620.

Bishop, C.V. and F. Stormshak. 2008. Minireview. Non-genomic actions of progesterone and estrogens in regulating reproductive events in domestic animals. Vet. J. 176:270-280.

Stormshak, F. and C.V. Bishop. 2008. Board-invited review: Estrogen and progesterone signaling: genomic and nongenomic action in domestic ruminants. J. Anim. Sci. 86:299-315.

Roselli, C.E., S. Bocklandt, H.L. Stadelman, T. Wadsworth, E. Vilain and F. Stormshak. 2008. Prolactin expression in the sheep brain. Neuroendocrinology 87:206-215.

Stormshak, F., C.T. Estill, J.A. Resko and C.E. Roselli. 2008. Changes in LH secretion in response to an estradiol challenge in male- and female-oriented rams and ewes. Reproduction 135:733-738.

Bishop, C.V., T. Filtz, Y. Zhang, O. Slayden and F. Stormshak. 2008. Progesterone suppresses an oxytocin-stimulated signal pathway in COS-7 cells transfected with the oxytocin receptor. Steroids 73:1367-1374.

Roselli, C.E., H. Stadelman, R. Reeve, C.V. Bishop and F. Stormshak. 2007. The ovine sexually dimorphic nucleus of the medial preoptic area is organized prenatally by testosterone. Endocrinology 148:4450-4457.

Bishop, C.V. and F. Stormshak. 2006. Nongenomic action of progesterone inhibits oxytocin-induced phosphoinositide hydrolysis and prostaglandin F2a secretion in the ovine endometrium. Endocrinology 147:937-942.

Roselli, C.E., J.M. Schrunk, H.L. Stadelman, J.A. Resko and F. Stormshak. 2006. The effect of aromatase inhibition on the sexual differentiation of the sheep brain. Endocrine 29:501-511.

Roselli, C.E., J.A. Resko and F. Stormshak. 2006. Expression of steroid hormone receptors in the fetal sheep brain during the critical period of sexual differentiation. Brain. Res. 1110:76-80.

Burke, J.M., C. Bishop and F. Stormshak. 2006. Reproductive characteristics of endophyte-infected or novel tall fescue fed ewes. Livestock Science 104:103-111.

Waite, A.L., D.W. Holtan and F. Stormshak. 2005. Changes in bovine luteal progesterone metabolism in response to exogenous prostaglandin F2a. Dom. Anim. Endocrinol. 28:162-171.

Dunlap, K.A. and F. Stormshak. 2004. Nongenomic inhibition of oxytocin binding by progesterone in the ovine uterus. Biol. Reprod. 70:65-69.

Roselli, C.E., K. Larkin, J.A. Resko, J.N. Stellflug and F. Stormshak. 2004. The volume of a sexually dimorphic nucleus in the ovine medial preoptic area/anterior hypothalamus varies with sexual partner preference. Endocrinology 145:478-483.

Roselli, C.E., K. Larkin, J.M. Schrunk and F. Stormshak. 2004. Sexual partner preference, hypothalamic morphology and aromatase in rams. Physiol. Behav. 83:233-245.

Paslay, E.M., J.R. Jaeger, U. Salli and F. Stormshak. 2003. Ovarian function in ewes after treatment with mifepristone early during the oestrous cycle. Reproduction 125:205-210.

Roselli, C.E., J.A. Resko and F. Stormshak. 2003. Estrogen synthesis in fetal sheep brain: Effect of maternal treatment with an aromatase inhibitor. Biol. Reprod. 68:370-374.

Stormshak, F. 2003. Biochemical and endocrine aspects of oxytocin production by the mammalian corpus luteum. Reprod. Biol. Endocrinol. 1:92. (Online BioMed Central)

Salli, U., N. Saito and F. Stormshak. 2003. Spatiotemporal interactions of myristoylated alanine-rich C kinase substrate (MARCKS) protein with the actin cytoskeleton and exocytosis of oxytocin upon prostaglandin F2a stimulation of bovine luteal cells. Biol. Reprod. 2053-2058.

Roselli, C.E., J.A. Resko and F. Stormshak. 2002. Hormonal influences on sexual partner preference in rams. Arch. Sexual Behav. 31:43-49.

Roselli, C.E., F. Stormshak, J.N. Stellflug and J.A. Resko. 2002. Relationship of serum testosterone concentrations to mate preference in rams. Biol. Reprod. 67:263-268.

Hu, J., T.E. Ludwig, U. Salli, F. Stormshak and M.A. Mirando. 2001. Autocrine/paracrine action of oxytocin in pig endometrium. Biol. Reprod. 64:1682-1688.

Salli, U. and F. Stormshak. 2001. Prostaglandin F2a-activated protein kinase Ca phosphorylates myristoylated alanine-rich C kinase substrate protein in bovine luteal cells.Endocrine 16:83-88.

Rueda, B.R., I.R. Hendry, W. J. Hendry III, F. Stormshak, O.D. Slayden and J.S. Davis. 2000.  Decreased progesterone levels and progesterone receptor antagonists promote apoptotic cell death in bovine luteal cells.  Biol. Reprod. 62:269-276.

Pinckard, K.L., J. Stellflug, J.A. Resko, C.E. Roselli and F. Stormshak. 2000. Review: Brain aromatization and other factors affecting male reproductive behavior with emphasis on the sexual orientation of rams.  Dom. Anim. Endocrinol.18:83-96.

Filley, S.J., H.A. Turner and F. Stormshak. 2000.  Plasma fatty acids, prostaglandin F2a metabolite, and reproductive response in postpartum heifers fed rumen bypass fat.  J. Anim. Sci. 78:139-144.

Pinckard, K.L., J. Stellflug and F. Stormshak. 2000. Influence of castration and estrogen replacement on sexual behavior of female-oriented, male-oriented, and asexual rams.  J. Anim. Sci.  78:1947-1953.

Salli, U., S. Supancic and F. Stormshak. 2000.  Minireview.  Phosphorylation of myristoylated alanine rich C kinase substrate (MARCKS) protein is associated with bovine luteal oxytocin exocytosis.  Biol. Reprod.  63:12-20.

Roselli, C.E., F. Stormshak and J.A. Resko. 2000. Distribution of aromatase mRNA in the ram hypothalamus: an in situ hybridization study.  J. Neuroendocrinol.  7:656-664.

Filley, S., S. Supancic, U. Salli, K. Orwig and F. Stormshak. 2000. Myristoylated alanine-rich C kinase substrate protein and mRNA in bovine corpus luteum during the estrous cycle.  Endocrine12:289-294.